Professor Emeritus, Biophysics; PhD, University of Chicago
As an assistant professor of biology at Harvard University, he taught physiology in an introductory biology course, Nat. Sci. 5, led by George Wald and pursued research on photoreceptors, the rods and cones of our eyes. In 1969, he chose to join the Biophysics Department at Johns Hopkins, where his research on the rapid diffusion of rhodopsin in photoreceptors led to the Cole award from the Biophysical Society.